QBARS - v14n1 The Parentage of Hybrid Azaleas

The Parentage of Hybrid Azaleas
by Clark D. Paris
Department of Horticulture
Michigan State University*

*The author wishes to thank Prof. C. E. Lewis for his assistance in collecting this data.

In 1753 when the immortal Linnaeus described the genera Azalea and Rhododendron , he initiated a controversy which has continued for almost two centuries. Most taxonomists agree that any line of demarcation between these two genera is entirely arbitrary; yet in 1942, Lemmon and McKay described a new species as Azalea furbishii. Shakespeare said that "a rose by any other name would smell as sweet". In this paper all species of "azaleas" are considered of the genus Rhododendron .

Regardless of the validity of the genus Azalea , this name has been retained for a group of plants that are very popular wherever they can be grown successfully. The wild species have been cultivated since the seventeenth century. When the pioneer hybridizers began crossing species of plants, the azaleas were among the first with which they worked. As more rhododendrons were introduced, these were crossed with the existing varieties. Each breeder developed his own breeding objectives and worked towards them until definite races of hybrids became known. Usually these races were named for the originator or the place where he worked. A complete list of azalea hybrid groups, as well as certain pertinent data about each, is given in Table I. The introduction dates of the hybrids were obtained directly from the originators, or their descendents. A study of the parentage of hybrid azaleas has revealed that, as far as is known, only twenty-seven species have been used. These, with the native habitat, color, and year of introduction, are given in Table II. The nomenclature is that given by Lee (4). Rehder (6) relegates some of these to a varietal status. No attempt is being made to give the pros and cons of this debate.

The main purpose of this paper is to present a chart to show the parentage of the introduced hybrid azaleas. The horizontal line above the name of each hybrid connects with a vertical line from each species or hybrid that was used in the production of that particular group. Any one variety does not necessarily contain blood from all parents. A broken line is used if the parent is in doubt. The Southern Indian hybrids and double Ghent hybrids are selections from Belgian Indian hybrids and Ghent hybrids, respectively. The Shammarello hybrids are inbred selections of Mollis hybrids.

This chart is just as accurate as the sources of information used. Like all progeny or pedigree records the originator's information is accepted as prima-facie evidence. Although it is probably at a bare minimum the possibility of deliberate misinformation cannot be wholly discounted. Several other sources of error are possible. One of the more plausible is the misidentification of parent material. Most people do not systematically check the material with which they are working, and then too, the same name has sometimes been applied to different plants. The old practice of giving Latin names to hybrids adds to the confusion. Sometimes the originator did not keep adequate records and depended on memory. Another early practice was the mixing of pollen and then when something of value was obtained an attempt was made to postulate the pollen parent by studying the hybrid. A less obvious pitfall is apogamy. When the seed is produced without actual fertilization occurring. The seeds so produced actually contain only maternal tissue.

Table I
Introduced Hybrid Azalea Groups

Hybrid Group

Originator

Introduced

Abbott

Frank L. Abbott, Bellows Falls, Vermont

About 1943

Albican

Anthony Waterer, Woking, Surrey, England

About 1870

Allan*

Walter Allan, Summerville, N. Carolina

1950

Amoena

Various English growers

1860

Arends

George Arends, Ronsdorf, Germany

1926

Arnold

Jackson T. Dawson, Jamaica Plain, Mass.

About 1910

Belgian Indian

Growers in England and Belgium

Early 1800's

Blaauw*

J. Blaauw and Co., Boskoop, Holland


Bobbink and Atkins
Macrantha

Bobbink and Atkins, East Rutherford, N.J.

About 1942

Chisholm-Merritt

Julian Chisholm, Garrett Park, Maryland

About 1947

Coe

Frederick W. Coe, Bethesda, Maryland


Coolidge

Coolidge Rare Plant Gardens, Pasadena, Calif.

1930's

Dawson

Henry S. Dawson, Eastern Nurseries. Holleston, Mass.

After 1923

De Wilde

Roland de Wilde, Rhodo-Lake Nurseries, Bridgeton, N. J.

1947

Deerfield

Deerfield Nursery, Deerfield Street P.O., N.J.

1950

Dosser*

Lillie Dosscr, Centralia, Washington

1955

Exbury Evergreen

Lionel de Rothschild, Exbury, Southampton, England

About 1955

Feldyk

Felix and Dijhuis, Boskoop, Holland

1930

Felix and Dijhuis

Felix and Dijhuis, Boskoop, Holland

1946

Ferndown*

D. Stewart and Son, Ltd., Ferndown, Dorset, England

1946

Gable

Joseph B. Gable, Stewartstown, Pa.

1927

Ghent

Growers around Ghent, Belgium

Early 1830's

Girard

Peter E. Girard, Girard Bros. Nursery, Geneva, Ohio

1954

Glenn Dale

United States Department of Agriculture

1941

Hage*

W. C. Hage and Co., Boskoop, Holland

1938

Henny

Rudolph Henny, Brooks, Oregon

Not Introduced

Hirado

Unknown. Island of Hirado, Japan

1952

Kaempferi
(Malvatica)

P. M. Koster, Boskoop, Holland

1920's

Kingsville*

Henry J. Hohman, Kingsville Nurseries, Kingsville, Maryland


Knaphill

Anthony Waterer, Woking, Surrey, England

About 1870

Knaphill Strain

Knap Hill Nursery, Woking, Surrey, England

1860's

Slocock Strain

Goldsworth Old Nursery, Ltd., Woking

1947

Ilam Strain

Edgar Stead, Ilam Estate, Christchurch, New Zealand


Exbury Strain

Lionel de Rothschild, Exbury, Southampton, England

Early 1930's

Koppeschaar*

W. F. Koppeschaar, Boskoop, Holland


Kurumanthum

K. Wada, Kakoneya Nurseries, Numazushi, Japan

Before 1941

Kurume

Mitozo Sakamoto, Kurume, Japan

Early 1800's

Lackamas

Ben Lancaster, Lackamas Gardens, Camas, Washington

1951

Macrindicum

K. Wada, Kakoneya Nurseries, Numazushi, Japan

Before 1941

Mayo

R. P. Mayo Nurseries, Augusta Georgia

About 1940

Mollis

Louis Van Houtte, Ghent, Belgium

Before 1873

Mortier

P. Mortier, Ghent, Belgium

About 1830

Occidentale

Growers in England and Holland

About 1895

Ornatum

J. R. Gowen, Highclere, England

About 1831

Ouden

H. den Ouden and Son, Boskoop, Holland

1938

Pericat

Alphone Pericat, Collingdale, Pa.

1931

Petry

Mrs. Frank Petry, Prichard, Alabama

After 1948

Pride

Orlando S. Pride, Butler, Pennsylvania

After 1934

Puyallup Valley

Leonard F. Frisbie, Puyallup, Washington

About 1954

Rustica Fiore Pleno
(Mixtum)

Charles Vuylsteke, Belgium

About 1900

Rutherford

Bobbink & Atkins. East Rutherford, N.J.

1920's

Sander

Charles Sander, Brookline, Mass.

About 1885

Satsuki

Unknown, Japan

Before 1700

Sawada

K. Sawada, Overlook Nurseries, Mobile, Ala.

About 1950

Scabrume

K. Wada, Kakoneya Nurseries, Numazushi, Japan

Before 1941

Shammarello

A. Shammarello, South Euclid, Ohio

Not Introduced

Sherwood

Sherwood Nursery Co., Corbett, Oregon

1935

Southern Indian

Magnolia Gardens, Charleston, S. Carolina

1840

Ten Oaks

Ten Oaks Nursery, Clarksville, Maryland

1948

Viscosepalum

Growers in England

Early 1800's

Vuyk

Vuyk van Nes Nursery, Boskoop, Holland

1926

Wadai

K. Wada, Kakoneya Nurseries, Numazushi, Japan

Before 1941

Westfall

Quincy R. Westfall, California

About 1950

Yerkes-Pryor

Guy E. Yerkes, U. S. D. A., Beltsville. Maryland


*Parentage unknown.

Table II
Rhododendron Species Used In The Production Of Hybrid Azaleas

Species

Native Habitat

Introduced

Color

R. arborescens (Pursh) Torr.

Eastern U. S.

Before 1814

White or pinkish

R. atlanticum (Ashe) Rehd.

Eastern U. S.

About 1916

White or pinkish

R. bakeri Lemm. & McKay

Southeastern U. S.

1938

Deep yellow to orange scarlet

R. calendulaceum (Michx.) Torr.

Eastern U. S.

Before 1800

Yellow to scarlet

R. canescens (Michx.) Sweet

Southern U. S.

1750?

Pink and white

R. indicum (L.) Sweet

Japan

Before 1680

Bright red to scarlet

R. japonicum (Gray) Suringar

Japan

1861

Orange-red or salmon-red to brick-red

R. kaempferi Planch.

Japan

1892

Red or orange-red to pink

R. kiusianum Mak.

Japan

1955

Purple

R. luteum Sweet

Asia Minor, Eastern Europe

1792

Yellow

R. macropetalum Maxim.

Japan

1863

Rose-lilac to rose-purple

R. molle (B1.) G. Don

China

1823

Golden-yellow

R. mucronatum G. Don

Japan

1819

White

R. nudiflorum (L.) Torr.

Eastern U. S.

About 1930

Light pink or white

R. obtusum (Lindl.) Planch.

Japan

About 1844

Orange-red to bright red

R. occidentale Gray

Western U. S.

1850

White or pinkish

R. oldhamii Maxim.

Formosa

1878

Red

R. phoenicum G. Don

Unknown, may be hybrid

1824

Rose-purple to magenta

R. poukhanense Levl.

Korea

1905

Pale lilac-purple

R. ripense Mak.

Japan

?

Rose-purple

R. roseum (Loisel.) Rehd.

Eastern U. S.

About 1790

Bright pink

R. scabrum G. Don

Japan

Before 1911

Rose-red to scarlet

R. schlippenbachii Maxim.

Eastern Asia

1893

Pale to rosy pink

R. simsii Planch.

China, Formosa

1808

Rose-red to bright or dark red

R. speciosum Sweet

Georgia, U.S.A.

1830

Scarlet-red

R. tschonoskii Maxim.

Japan, Korea

1878

White

R. viscosum (L.) Torr.

Eastern U. S.

Before 1731

White or suffused pink

Fig06.jpg

Fig. 6

BIBLIOGRAPHY

1. Arends, Georg, Mein Leben als Gartner and Zuchter, Stuttgart: Eugen Ulmer, 1951.

2. Bowers, Clement Gray, Rhododendrons and Azaleas, New York: Macmillan, 1936.

3. Hume, H. Harold, Azaleas, New York: Macmillan, 1956.

4. Lee, Frederic P., The Azalea Book, Princeton: D. Van Nostrand, 1958.

5. Pinckney, G. H., Hardy Hybrid Azaleas-Their Evolution and Range, Royal Horticultural Society, The Rhododendron and Camellia Year Book, 9:31-36, 1955.

6. Rehder, Alfred, Manual of Cultivated Trees and Shrubs, Second edition, New York: Macmillan, 1940.

7. Wilson, E. H. and Rehder, Alfred, A Monograph of Azaleas, Cambridge, Harvard Univ. Press, 1921.

8. Wyman, Donald, Shrubs and Vines for American Gardens, New York: Macmillan, 1949.